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Elephants have a good memory and they can recall the smell of a relative even after a decade apart. 

When Franziska Horner and her team 1 planned reunions (团聚) between two mother-daughter pairs at zoos in Germany, they took advantage of the chance to test the elephants' 2 . One pair had been 3 for two years, 4 the other had spent 12 years apart.

Franziska 5  faecal (排泄物的) samples from these African elephants and others at German zoos, carrying the smelly 10-to-15-kilogram faeces around in her tiny car. Her team 6  these samples one at a time to the four elephants 7  their reunions. 

When they encountered faeces from 8  elephants, either those in the same zoo or unfamiliar animals, they sniffed (闻) the faeces piles and walked away. But when presented with a 9  from the mother or daughter they were due to be reunited with, the female elephants 10  sniffed the samples and showed a variety of 11  , from making deep sounds to swinging their ears. 

Such reactions may be linked to positive emotions, the researchers say. "That was amazing and intense," says Franziska. "We were sure they did 12 , and they knew exactly what they were 13 there. The sample size was small, but it would be 14 to do similar tests on elephants that weren't being reunited. "

"I am not surprised that elephants have memories, especially in social contexts, that 15 a long time," says another scientist.

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